So far we’ve studied only the evolution of a single trait, e.g., height or weight. But organisms have many traits, and they evolve at the same time. How can we understand their simultaneous evolution? The basic framework of the quantitative genetic approach was first outlined by Russ Lande and Steve Arnold [2]. Let z1, z2, . . . , zn be the phenotype of each character that we are studying. We’l...