A new Middle Jurassic caddisfly (Trichoptera, Hydrobiosidae) from China

The order Trichoptera, so-called “hairy wings”, contains three suborders, namely Annulipalpia, Integripalpia, and Spicipalpia, with over 13,000 known extant species. The order is widely distributed around the world, except for the polar regions (Morse 2012), and has an extensive fossil record, with 37 families, 193 genera, and more than 660 species reported to date (Handlirsch 1906–1908, 1939; Meunier 1918; Martynova 1958; Sukastcheva 1968, 1973, 1982, 1990; Erickson 1983; Novokshonov 1993; Botosaneanu 1995; Jarzembowski 1995; Novokshonov et al. 1995; Ansorge 2002; Ivanov & Sukatsheva 2002; Ivanov & Melnitsky 2005; Wichard 2007; Wichard et al. 2009, 2011). In particular, four families of Trichoptera (including nine genera and 15 species) have been documented from China (Hong 1983; Lin 1986; Ren et al. 1995; Wang et al. 2009a, 2009b; Davis et al. 2010). Among caddisflies, the Spicipalpia consists of five extant families, namely Rhyacophilidae, Hydrobiosidae, Glossosomatidae, Hydroptilidae and Ptilocolepidae (Ross 1956; Schmid 1970; Morse 1997; Malicky 2001, 2005; Ward et al. 2004). The family Hydrobiosidae was first erected by Ulmer (1905) as a subfamily of Rhyacophilidae, and granted with family rank by Schmid (1989). This is a large family of approximately 50 genera (Holzenthal et al. 2007). So far, only three genera and three species of the fossil Hydrobiosidae have been reported (Table 1).