Four new species of Anatomidae (Mollusca: Vetigastropoda) from the Indian Ocean (Reunion, Mayotte) and Australia, with notes on a novel radular type for the family

Four new species of Anatomidae are described: Anatoma herberti n. sp. with strong axial sculpture on the shoulder and base; A. austrolissa n. sp. with almost smooth sculpture except for axial cords in the adumbilical half of the base; A. boucheti n. sp. with sunken protoconch and selenizone that starts after more than one teleoconch I whorl; and A. fl exidentata n. sp. with a highly modifi ed radula, shared only with A. austrolissa, among known anatomid species. Three of the species are only known from the Indian Ocean, while the more deep-water A. austrolissa is known from Reunion Island and New South Wales, Australia. The radula of A. fl exidentata and A. austrolissa is strikingly different from that of other Anatomidae and Vetigastropoda in that it has fl exible equally-shaped teeth in the central fi eld and fi lamentous teeth in the marginal fi eld. Similar radular morphologies are known from Calliostomatidae.