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7 Centre for Invasion Biology, Department of Conservation Ecology and Entomology, Stellenbosch 8 University, Matieland 7602, South Africa; Department of Biology, Juniata College, Huntingdon, 9 Pennsylvania 16652, USA; Department of Electrical and Electronic Engineering, Stellenbosch 10 University, Matieland 7602, South Africa; Centre for Invasion Biology, Department of Botany and 11 Zoology, Stellenbosch University, Matieland 7602, South Africa. 12 13 *Author for correspondence ([email protected]) 14 http://jeb.biologists.org/lookup/doi/10.1242/jeb.095554 Access the most recent version at J Exp Biol Advance Online Articles. First posted online on 30 August 2013 as doi:10.1242/jeb.095554