new locality data are given for eight species of geotrupidae and 65 species and sub-species of scarabaeidae dung beetles from iran. the species onthophagus (palaeonthophagus) anatolicus petrovitz, 1962, and onitis alexis ssp. septentrionalis balthasar, 1942, are recorded for the first time from iran.

The external and internal male genitalia of 327 species of 11 tribes of the subfamily Scarabaeinae, including species of Deltochilini, Scarabaeini, Gymnopleurini, Ateuchini, and Coprini, among others, were examined. Descriptions of the variations in the genital segment, the aedeagus, the internal sac, and its sclerites and raspules are presented. An exhaustive comparison of structures, names, a...

The dung beetles (Scarabaeinae) include ca. 5000 species and exhibit a diverse array of morphologies and behaviors. This variation presumably reflects the adaptation to a diversity of food types and the different strategies used to avoid competition for vertebrate dung, which is the primary breeding environment for most species. The current classification gives great weight to the major behavio...