Corresponding author. Mailing address: Department of Veterinary Microbiology and Pathology, 17 Washington State University College of Veterinary Medicine, P. O. Box 647040, Pullman, WA 18 99164-7040. Phone: (509) 335-6075. Fax: (509) 335-8529. E-mail: [email protected]. 19 AEM Accepts, published online ahead of print on 6 September 2013 Appl. Environ. Microbiol. doi:10.1128/AEM.02173-13 Co...