HE gene A , in maize was first reported, with its mutant allele a,, by EMERSON (1918, 1921). In these reports and later publications by EMERSON and ANDERSON (1932) and LAUGHNAN (1948) the A , allele found in North American races was shown to be necessary for the production of anthocyanin pigments in various tissues of the corn plant. With the complementary genes A,, B, Pl, and R' or r', A , pro...