24 25 The combined effects of isolation within an unfavourable matrix and vegetation 26 development were studied in patches of scrub vegetation on a tropical inselberg (Nouragues, 27 French Guiana). Three thickets ~10 m 2 area were sampled, located at ~30 m distance from 28 each other, in which the same vegetation types were present at less than 3 m distance: 29 canopies of Clusia minor (Clusia...