Cretaceous ichthyosaurs have typically been considered a small, homogeneous assemblage sharing a common Late Jurassic ancestor. Their low diversity and disparity have been interpreted as indicative of a decline leading to their Cenomanian extinction. We describe the first post-Triassic ichthyosaur from the Middle East, Malawania anachronus gen. et sp. nov. from the Early Cretaceous of Iraq, and...

Australian dinosaurs have played a rare but controversial role in the debate surrounding the effect of Gondwanan break-up on Cretaceous dinosaur distribution. Major spatiotemporal gaps in the Gondwanan Cretaceous fossil record, coupled with taxon incompleteness, have hindered research on this effect, especially in Australia. Here we report on two new sauropod specimens from the early Late Creta...

Plesiosaurs were the longest-surviving group of secondarily marine tetrapods, comparable in diversity to today's cetaceans. During their long evolutionary history, which spanned the Jurassic and the Cretaceous (201 to 66 Ma), plesiosaurs repeatedly evolved long- and short-necked body plans [1, 2]. Despite this postcranial plasticity, short-necked plesiosaur clades have traditionally been regard...

The first flowering plant fossils occur as rare, undiverse pollen grains in the Early Cretaceous (Valanginian-Hauterivian). Angiosperms diversified slowly during the Barremian-Aptian but rapidly during the Albian-Cenomanian. By the end of the Cretaceous, at least half of the living angiosperm orders were present, and angiosperms were greater than 70% of terrestrial plant species globally. The r...

The flowering plants that dominate modern vegetation possess leaf gas exchange potentials that far exceed those of all other living or extinct plants. The great divide in maximal ability to exchange CO(2) for water between leaves of nonangiosperms and angiosperms forms the mechanistic foundation for speculation about how angiosperms drove sweeping ecological and biogeochemical change during the...

Enantiornithes are the most successful clade of Mesozoic birds. Here, we describe a new enantiornithine bird, Cruralispennia multidonta gen. et sp. nov., from the Protopteryx-horizon of the Early Cretaceous Huajiying Formation of China. Despite being among the oldest known enantiornithines, Cruralispennia displays derived morphologies that are unexpected at such an early stage in the evolution ...

We here describe a new Early Cretaceous (early Albian) eutherian mammal, Sasayamamylos kawaii gen. et sp. nov., from the 'Lower Formation' of the Sasayama Group, Hyogo Prefecture, Japan. Sasayamamylos kawaii is characterized by a robust dentary, a distinct angle on the ventral margin of the dentary at the posterior end of the mandibular symphysis, a lower dental formula of 3-4 : 1 : 4 : 3, a ro...

Recent discoveries of basal dromaeosaurids from the Early Cretaceous Jiufotang and Yixian formations of Liaoning, China, add significant new information about the transition from non-avian dinosaurs to avians. Here we report on a new dromaeosaurid, Tianyuraptor ostromi gen. et sp. nov., from the Early Cretaceous Yixian Formation of western Liaoning, China, based on a nearly complete skeleton. T...

The Tirgan Formation (Barremian – Aptian) is exposed in the Kopet Dagh in northeast Iran. One stratigraphic section in Zavin was measured with a thickness of 110 meters. This Formation in section consists of three parts (including lower carbonate, limy shale – marl and upper carbonate rocks). Based on the study of 94 thin sections, 10 carbonate and 2 siliciclastic lithofacies have been identifi...

The fossil record of Australian dinosaurs in general, and theropods in particular, is extremely sparse. Here we describe an ulna from the Early Cretaceous Eumeralla Formation of Australia that shares unique autapomorphies with the South American theropod Megaraptor. We also present evidence for the spinosauroid affinities of Megaraptor. This ulna represents the first Australian non-avian therop...