'Environmental Sciences Division, Oak Ridge National Luboratory, P.O. Box 2008, Oak Ridge, Tennessee 37831 USA LDeparrment of Natural Resources and Environmental Sciences, University of Illinois, Urbana, Illinois 61801 USA ?Institute of Ecology. University of Georgia. Athens, Georgia 30602 USA -'Ecosystems Center. Marine Biological Luboratory, Woods Hole, Massachusetts 02543 USA 'Department of ...