J. M. H. Knops,* K. L. Bradley and D. A. Wedin School of Biological Sciences, School of Natural Resource Sciences, University of Nebraska, 348 Manter Hall, Lincoln, NE 68588, USA. *Correspondence: E-mail: jknops2@unl.edu Abstract Plant species are hypothesized to impact ecosystem nitrogen cycling in two distinctly different ways. First, differences in nitrogen use efficiency can lead to positiv...