Groundnut (Arachis hypogaea) plants with rosette disease contain a manually transmissible virus, groundnut rosette virus (GRV), which depends on a luteovirus, groundnut rosette assistor virus (GRAV), for transmission by the aphid Aphis craccivora. No virus-like particles have been reported for GRV but infected plants yield infective ssRNA. Infected leaves also contain dsRNA with prominent elect...