l. b. silvestro laboratory of functional biology and biotechnology (biolab-inbiotec-conicet-cic), national university of the center of the province of buenos aires, argentina. f. biganzoli department of quantitative methods and information systems, faculty of agronomy, university of buenos aires, argentina. h. forjn barrow-inta experimental station, tres arroyos, province of buenos aires, argentina. a. albanesi chair in microbiology and ecology, agricultural faya national university of santiago del estero, argentina. a. m. arambarri spegazzini institute, fcnym, national university of la plata, argentina. l. manso barrow-inta experimental station, tres arroyos, province of buenos aires, argentina. m. v. moreno