hamid ghasemi eastern azerbaijan agriculture and natural resources research center, p.o. box: 53555-141 tabriz, iran. arash jouladeh roudbar department of fisheries, faculty of natural resources, sari university of agricultural sciences and natural resources, sari, iran. soheil eagderi department of fisheries, faculty of natural resources, university of tehran, karaj, iran. keivan abbasi aquatic ecology department of the inland water aquaculture research center, p.o. box 66, bandar anzali, iran. saber vatandoust department of fisheries, babol branch of islamic azad university, mazandaran, iran. hamid reza esmaeili department of biology, college of sciences, shiraz university, shiraz, 71454–iran.