Research Article| December 29, 2022 DEEP TIME BIOGEOMORPHOLOGY 2: ANIMALS AS ANCIENT ECOSYSTEM ENGINEERS ANTHONY P. SHILLITO; SHILLITO 1Department of Geological Sciences, University Saskatchewan, 114 Science Place, Campus Drive, Saskatoon SK S7N 5E2, Canada Search for other works by this author on: GSW Google Scholar NEIL S. DAVIES; DAVIES 2Department Earth Cambridge, Downing Street, Cambridge ...