The Oligocene (Rupelian) Byram Formation (Vicksburg Group) in Alabama, USA, is divided into three members, including (in ascending order) the Glendon Limestone, unnamed marl, and Bucatunna Clay. The marine units Alabama have been historically under-investigated, but bulk samples recently obtained from Limestone Member exposures at site AWa-9 Washington County yielded 20 unequivocal elasmobranch...