8 MRC University of Glasgow Centre for Virus Research, Institute of Infection, 9 Immunity and Inflammation, College of Medical, Veterinary and Life Sciences, 10 University of Glasgow, United Kingdom 11 EPHE, Université de Lyon 1, INRA, UMR754, École Nationale Vétérinaire de Lyon, 12 IFR 128, Lyon, France 13 Istituto G. Caporale, Teramo, Italy 14 ISPAAM, National Research Council (CNR), Naples, ...