The extinct Devonian placoderms (armoured jawed fishes) [1,2] are central to the question of tooth origins, because some have denticulate ‘toothplates’ within the mouth cavity. A key question is whether these gnathal plates were modified from external dermal bones, or had ‘denticles’ representing true teeth with pulp cavities [3, fig. 2h]. The recent contribution by Rücklin & Donoghue [4] confu...