The carbohydrate moieties of arabinogalactan proteins (AGPs) are essential for their physiological functions and undergo rapid turnover in vivo. Degradation of the carbohydrate moieties of AGPs seems to occur by concerted action of several glycosidases, among them a-L-arabinofuranosidase, b-Dgalactosidase, and b-D-glucuronidase. Here, a bifunctional a-L-arabinofuranosidase/b-D-xylosidase from i...