Vegetation succession was studied for 12 years on an abandoned nutrient-poor small field surrounded by a strip of natural grassland. No fertilizers or herbicides were applied. Few weeds or annuals were present during this period. Only two plants, i.e. Agrostis capillaris and Festuca rubra , dominated during the 12 years. At two spatial scales (0.04 and 0.4 m) a dramatic increase in species rich...